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What is a spiculum?

What is a spiculum? Here are some definitions.

Noun
  1. A thrusting javelin used by Romans that replaced the pilum in the late 3rd century.
  2. A sharp, pointed crystal, especially of ice.
  3. (zoology) A sharp, needle-like structure, especially those making up the skeleton of a sponge.
  4. (astronomy) A small radial emission of gas seen in the chromosphere and corona of the sun.
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This surrounds the cloacal outlet, the latter concealing a double spiculum.

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