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What is a groundling?

What is a groundling? Here are some definitions.

Noun
  1. Any of various plants or animals living on or near the ground, as a benthic fish or bottom feeder, especially:
    1. The spined loach (Cobitis taenia), weather loach ( Misgurnus fossilis), or other member of the loaches.
    2. The ringed plover, Charadrius hiaticula.
  2. An audience member in the cheap section (usually standing; originally in Elizabethan theater).
  3. By extension, an individual of uncultivated or uncultured taste.
  4. One who is confined to the ground, especially:
    1. (military, slang) A soldier who fights on the ground or serves as ground crew, as opposed to a seaman, pilot, etc.
    2. (fantasy) A member of a race that lives primarily underground, such as a dwarf.
  5. (Abrahamic religions) Adam, before eating the apple of knowledge of good and evil (emphasizing his creation from the ground).
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Examples
Pumpkins are among the most imposing of all groundling growths.
Fertilized groundling and sturgeon eggs were irradiated at stages from fertilization to middle gastrulation.
The two of them were naked, the groundling and his woman, they were not ashamed.
But the gentleman, now coming up the drive, was not in the proper frame of mind for groundling observation.
Simon noticed a groundling look intently at Walsingham's purse and nudge his mate.
It was obvious that the brown-haired groundling had been involved in some heavy hand-to-hand combat recently.

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